Pleistocene

Pleistocene
Pleistocene
2.58 – 0.0117 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Map of the world during the Last Glacial Maximum
Chronology
←	Meghalayan; Northgrippian; Greenlandian
	Subdivision of the Quaternary according to the ICS, as of 2021.; Vertical axis scale: millions of years ago.;
Etymology
Name formality	Formal
Usage information
Celestial body	Earth
Regional usage	Global (ICS)
Definition
Chronological unit	Epoch
Stratigraphic unit	Series
Time span formality	Formal
Lower boundary definition	Base of magnetic polarity chronozone C2r (Matuyama).; Extinction of the haptophytes Discoaster pentaradiatus and Discoaster surculus;
Lower boundary GSSP	Monte San Nicola Section, Gela, Sicily, Italy; 37.1469°N 14.2035°E
Lower GSSP ratified	2009 (as base of Quaternary and Pleistocene)
Upper boundary definition	End of the Younger Dryas stadial
Upper boundary GSSP	NGRIP2 ice core, Greenland; 75.1000°N 42.3200°W
Upper GSSP ratified	2008 (as base of Holocene)

The Pleistocene ( /ˈplaɪs.təsiːn, -toʊ-/ PLYSE-tə-seen, -⁠toh-,^[5] often referred to colloquially as the Ice Age) is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek πλεῖστος, pleīstos, 'most' and καινός, kainós (latinized as cænus), 'new'.

At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by the Isthmus of Panama, causing a faunal interchange between the two regions and changing ocean circulation patterns, with the onset of glaciation in the Northern Hemisphere occurring around 2.7 million years ago. During the Early Pleistocene (2.58–0.8 Ma), archaic humans of the genus Homo originated in Africa and spread throughout Afro-Eurasia. The end of the Early Pleistocene is marked by the Mid-Pleistocene Transition, with the cyclicity of glacial cycles changing from 41,000-year cycles to asymmetric 100,000-year cycles, making the climate variation more extreme. The Late Pleistocene witnessed the spread of modern humans outside of Africa as well as the extinction of all other human species. Humans also spread to the Australian continent and the Americas for the first time, co-incident with the extinction of most large bodied animals in these regions.

The aridification and cooling trends of the preceding Neogene were continued in the Pleistocene. The climate was strongly variable depending on the glacial cycle, with the sea levels being up to 120 metres lower than present at peak glaciation, allowing the connection of Asia and North America via Beringia and the covering of most of northern North America by the Laurentide Ice Sheet.

Etymology

Evolution of temperature in the Post-Glacial period at the very end of the Pleistocene, according to Greenland ice cores^[6]

Temperature rise marking the end of the Pleistocene, as derived from Antarctic ice core data.

Charles Lyell introduced the term "Pleistocene" in 1839 to describe strata in Sicily that had at least 70% of their molluscan fauna still living today. This distinguished it from the older Pliocene Epoch, which Lyell had originally thought to be the youngest fossil rock layer. He constructed the name "Pleistocene" ("most new" or "newest") from the Greek πλεῖστος (pleīstos, "most") and καινός (kainós (latinized as cænus), "new");^[7]^[8]^[9] this contrasts with the immediately preceding Pliocene ("newer", from πλείων (pleíōn, "more") and kainós) and the immediately subsequent Holocene ("wholly new" or "entirely new", from ὅλος (hólos, "whole") and kainós) epoch, which extends to the present time.

Dating

Hominin timeline

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Miocene

Pliocene

Pleistocene

Hominini

Nakalipithecus

Ouranopithecus

Oreopithecus

Sahelanthropus

Orrorin

Ardipithecus

Australopithecus

Homo habilis

Homo erectus

Homo bodoensis

Homo sapiens

Neanderthals,Denisovans

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Dispersal beyond Africa

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Earliest fire / cooking

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Earliest clothes

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Modern humans

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(million years ago)

The Pleistocene has been dated from 2.580 million (±0.005) to 11,650 years BP^[10] with the end date expressed in radiocarbon years as 10,000 carbon-14 years BP.^[11] It covers most of the latest period of repeated glaciation, up to and including the Younger Dryas cold spell. The end of the Younger Dryas has been dated to about 9640 BC (11,654 calendar years BP). The end of the Younger Dryas is the official start of the current Holocene Epoch. Although it is considered an epoch, the Holocene is not significantly different from previous interglacial intervals within the Pleistocene.^[12] In the ICS timescale, the Pleistocene is divided into four stages or ages, the Gelasian, Calabrian, Chibanian (previously the unofficial "Middle Pleistocene"), and Upper Pleistocene (unofficially the "Tarantian").^[13]^[14]^{[note 1]} In addition to these international subdivisions, various regional subdivisions are often used.

In 2009 the International Union of Geological Sciences (IUGS) confirmed a change in time period for the Pleistocene, changing the start date from 1.806 to 2.588 million years BP, and accepted the base of the Gelasian as the base of the Pleistocene, namely the base of the Monte San Nicola GSSP.^[16] The start date has now been rounded down to 2.580 million years BP.^[10] The IUGS has yet to approve a type section, Global Boundary Stratotype Section and Point (GSSP), for the upper Pleistocene/Holocene boundary (i.e. the upper boundary). The proposed section is the North Greenland Ice Core Project ice core 75° 06' N 42° 18' W.^[17] The lower boundary of the Pleistocene Series is formally defined magnetostratigraphically as the base of the Matuyama (C2r) chronozone, isotopic stage 103. Above this point there are notable extinctions of the calcareous nanofossils: Discoaster pentaradiatus and Discoaster surculus.^[18]^[19] The Pleistocene covers the recent period of repeated glaciations.

The name Plio-Pleistocene has, in the past, been used to mean the last ice age. Formerly, the boundary between the two epochs was drawn at the time when the foraminiferal species Hyalinea baltica first appeared in the marine section at La Castella, Calabria, Italy;^[20] however, the revised definition of the Quaternary, by pushing back the start date of the Pleistocene to 2.58 Ma, results in the inclusion of all the recent repeated glaciations within the Pleistocene.

Radiocarbon dating is considered to be inaccurate beyond around 50,000 years ago. Marine isotope stages (MIS) derived from Oxygen isotopes are often used for giving approximate dates.

Deposits

Pleistocene non-marine sediments are found primarily in fluvial deposits, lakebeds, slope and loess deposits as well as in the large amounts of material moved about by glaciers. Less common are cave deposits, travertines and volcanic deposits (lavas, ashes). Pleistocene marine deposits are found primarily in shallow marine basins mostly (but with important exceptions) in areas within a few tens of kilometres of the modern shoreline. In a few geologically active areas such as the Southern California coast, Pleistocene marine deposits may be found at elevations of several hundred metres.

Paleogeography and climate

The maximum extent of glacial ice in the north polar area during the Pleistocene Period

The modern continents were essentially at their present positions during the Pleistocene, the plates upon which they sit probably having moved no more than 100 km (62 mi) relative to each other since the beginning of the period. In glacial periods, the sea level would drop by over 100 m (330 ft) during peak glaciation, exposing large areas of present continental shelf as dry land.

According to Mark Lynas (through collected data), the Pleistocene's overall climate could be characterised as a continuous El Niño with trade winds in the south Pacific weakening or heading east, warm air rising near Peru, warm water spreading from the west Pacific and the Indian Ocean to the east Pacific, and other El Niño markers.^[21]

Glacial features[edit]

Pleistocene climate was marked by repeated glacial cycles in which continental glaciers pushed to the 40th parallel in some places. It is estimated that, at maximum glacial extent, 30% of the Earth's surface was covered by ice. In addition, a zone of permafrost stretched southward from the edge of the glacial sheet, a few hundred kilometres in North America, and several hundred in Eurasia. The mean annual temperature at the edge of the ice was −6 °C (21 °F); at the edge of the permafrost, 0 °C (32 °F).

Each glacial advance tied up huge volumes of water in continental ice sheets 1,500 to 3,000 metres (4,900–9,800 ft) thick, resulting in temporary sea-level drops of 100 metres (300 ft) or more over the entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions.

The effects of glaciation were global. Antarctica was ice-bound throughout the Pleistocene as well as the preceding Pliocene. The Andes were covered in the south by the Patagonian ice cap. There were glaciers in New Zealand and Tasmania. The current decaying glaciers of Mount Kenya, Mount Kilimanjaro, and the Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of Ethiopia and to the west in the Atlas mountains.

In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the east was covered by the Laurentide. The Fenno-Scandian ice sheet rested on northern Europe, including much of Great Britain; the Alpine ice sheet on the Alps. Scattered domes stretched across Siberia and the Arctic shelf. The northern seas were ice-covered.

South of the ice sheets large lakes accumulated because outlets were blocked and the cooler air slowed evaporation. When the Laurentide Ice Sheet retreated, north-central North America was totally covered by Lake Agassiz. Over a hundred basins, now dry or nearly so, were overflowing in the North American west. Lake Bonneville, for example, stood where Great Salt Lake now does. In Eurasia, large lakes developed as a result of the runoff from the glaciers. Rivers were larger, had a more copious flow, and were braided. African lakes were fuller, apparently from decreased evaporation. Deserts, on the other hand, were drier and more extensive. Rainfall was lower because of the decreases in oceanic and other evaporation.

It has been estimated that during the Pleistocene, the East Antarctic Ice Sheet thinned by at least 500 meters, and that thinning since the Last Glacial Maximum is less than 50 meters and probably started after ca 14 ka.^[22]

Major events

Ice ages as reflected in atmospheric CO₂, stored in bubbles from glacial ice of Antarctica

During the 2.5 million years of the Pleistocene, numerous cold phases called glacials (Quaternary ice age), or significant advances of continental ice sheets, in Europe and North America, occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter interglacials which lasted about 10,000–15,000 years. The last cold episode of the last glacial period ended about 10,000 years ago.^[23] Over 11 major glacial events have been identified, as well as many minor glacial events.^[24] A major glacial event is a general glacial excursion, termed a "glacial." Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials".

These events are defined differently in different regions of the glacial range, which have their own glacial history depending on latitude, terrain and climate. There is a general correspondence between glacials in different regions. Investigators often interchange the names if the glacial geology of a region is in the process of being defined. However, it is generally incorrect to apply the name of a glacial in one region to another.

For most of the 20th century only a few regions had been studied and the names were relatively few. Today the geologists of different nations are taking more of an interest in Pleistocene glaciology. As a consequence, the number of names is expanding rapidly and will continue to expand. Many of the advances and stadials remain unnamed. Also, the terrestrial evidence for some of them has been erased or obscured by larger ones, but evidence remains from the study of cyclical climate changes.

The glacials in the following tables show historical usages, are a simplification of a much more complex cycle of variation in climate and terrain, and are generally no longer used. These names have been abandoned in favour of numeric data because many of the correlations were found to be either inexact or incorrect and more than four major glacials have been recognised since the historical terminology was established.^[24]^[25]^[26]

Historical names of the "four major" glacials in four regions.
Region	Glacial 1	Glacial 2	Glacial 3	Glacial 4
Alps	Günz	Mindel	Riss	Würm
North Europe	Eburonian	Elsterian	Saalian	Weichselian
British Isles	Beestonian	Anglian	Wolstonian	Devensian
Midwest U.S.	Nebraskan	Kansan	Illinoian	Wisconsinan

Historical names of interglacials.
Region	Interglacial 1	Interglacial 2	Interglacial 3
Alps	Günz-Mindel	Mindel-Riss	Riss-Würm
North Europe	Waalian	Holsteinian	Eemian
British Isles	Cromerian	Hoxnian	Ipswichian
Midwest U.S.	Aftonian	Yarmouthian	Sangamonian

Corresponding to the terms glacial and interglacial, the terms pluvial and interpluvial are in use (Latin: pluvia, rain). A pluvial is a warmer period of increased rainfall; an interpluvial, of decreased rainfall. Formerly a pluvial was thought to correspond to a glacial in regions not iced, and in some cases it does. Rainfall is cyclical also. Pluvials and interpluvials are widespread.

There is no systematic correspondence of pluvials to glacials, however. Moreover, regional pluvials do not correspond to each other globally. For example, some have used the term "Riss pluvial" in Egyptian contexts. Any coincidence is an accident of regional factors. Only a few of the names for pluvials in restricted regions have been stratigraphically defined.

Palaeocycles

The sum of transient factors acting at the Earth's surface is cyclical: climate, ocean currents and other movements, wind currents, temperature, etc. The waveform response comes from the underlying cyclical motions of the planet, which eventually drag all the transients into harmony with them. The repeated glaciations of the Pleistocene were caused by the same factors.

The Mid-Pleistocene Transition, approximately one million years ago, saw a change from low-amplitude glacial cycles with a dominant periodicity of 41,000 years to asymmetric high-amplitude cycles dominated by a periodicity of 100,000 years.^[27]

However, a 2020 study concluded that ice age terminations might have been influenced by obliquity since the Mid-Pleistocene Transition, which caused stronger summers in the Northern Hemisphere.^[28]

Milankovitch cycles

Glaciation in the Pleistocene was a series of glacials and interglacials, stadials and interstadials, mirroring periodic changes in climate. The main factor at work in climate cycling is now believed to be Milankovitch cycles. These are periodic variations in regional and planetary solar radiation reaching the Earth caused by several repeating changes in the Earth's motion.

Milankovitch cycles cannot be the sole factor responsible for the variations in climate since they explain neither the long term cooling trend over the Plio-Pleistocene, nor the millennial variations in the Greenland Ice Cores. Milankovitch pacing seems to best explain glaciation events with periodicity of 100,000, 40,000, and 20,000 years. Such a pattern seems to fit the information on climate change found in oxygen isotope cores.

Oxygen isotope ratio cycles

In oxygen isotope ratio analysis, variations in the ratio of 18
O to 16
O (two isotopes of oxygen) by mass (measured by a mass spectrometer) present in the calcite of oceanic core samples is used as a diagnostic of ancient ocean temperature change and therefore of climate change. Cold oceans are richer in 18
O, which is included in the tests of the microorganisms (foraminifera) contributing the calcite.

A more recent version of the sampling process makes use of modern glacial ice cores. Although less rich in 18
O than sea water, the snow that fell on the glacier year by year nevertheless contained 18
O and 16
O in a ratio that depended on the mean annual temperature.

Temperature and climate change are cyclical when plotted on a graph of temperature versus time. Temperature coordinates are given in the form of a deviation from today's annual mean temperature, taken as zero. This sort of graph is based on another of isotope ratio versus time. Ratios are converted to a percentage difference from the ratio found in standard mean ocean water (SMOW).

The graph in either form appears as a waveform with overtones. One half of a period is a Marine isotopic stage (MIS). It indicates a glacial (below zero) or an interglacial (above zero). Overtones are stadials or interstadials.

According to this evidence, Earth experienced 102 MIS stages beginning at about 2.588 Ma BP in the Early Pleistocene Gelasian. Early Pleistocene stages were shallow and frequent. The latest were the most intense and most widely spaced.

By convention, stages are numbered from the Holocene, which is MIS1. Glacials receive an even number; interglacials, odd. The first major glacial was MIS2-4 at about 85–11 ka BP. The largest glacials were 2, 6, 12, and 16; the warmest interglacials, 1, 5, 9 and 11. For matching of MIS numbers to named stages, see under the articles for those names.

Fauna

Both marine and continental faunas were essentially modern but with many more large land mammals such as Mammoths, Mastodons, Diprotodon, Smilodon, tiger, lion, Aurochs, short-faced bears, giant sloths, Gigantopithecus and others. Isolated landmasses such as Australia, Madagascar, New Zealand and islands in the Pacific saw the evolution of large birds and even reptiles such as the Elephant bird, moa, Haast's eagle, Quinkana, Megalania and Meiolania.

The severe climatic changes during the Ice Age had major impacts on the fauna and flora. With each advance of the ice, large areas of the continents became totally depopulated, and plants and animals retreating southwards in front of the advancing glacier faced tremendous stress. The most severe stress resulted from drastic climatic changes, reduced living space, and curtailed food supply. A major extinction event of large mammals (megafauna), which included mammoths, mastodons, saber-toothed cats, glyptodons, the woolly rhinoceros, various giraffids, such as the Sivatherium; ground sloths, Irish elk, cave bears, Gomphotheres, dire wolves, and short-faced bears, began late in the Pleistocene and continued into the Holocene. Neanderthals also became extinct during this period. At the end of the last ice age, cold-blooded animals, smaller mammals like wood mice, migratory birds, and swifter animals like whitetail deer had replaced the megafauna and migrated north. Late Pleistocene bighorn sheep were more slender and had longer legs than their descendants today. Scientists believe that the change in predator fauna after the late Pleistocene extinctions resulted in a change of body shape as the species adapted for increased power rather than speed.^[29]

The extinctions hardly affected Africa but were especially severe in North America where native horses and camels were wiped out.

Various schemes for subdividing the Pleistocene

Asian land mammal ages (ALMA) include Zhoukoudianian, Nihewanian, and Yushean.
European land mammal ages (ELMA) include the Villafranchian, Galerian, and Aurelian
North American land mammal ages (NALMA) include Blancan (4.75–1.8), Irvingtonian (1.8–0.24) and Rancholabrean (0.24–0.01) in millions of years. The Blancan extends significantly back into the Pliocene.
South American land mammal ages (SALMA) include Uquian (2.5–1.5), Ensenadan (1.5–0.3) and Lujanian (0.3–0.01) in millions of years. The Uquian previously extended significantly back into the Pliocene, although the new definition places it entirely within the Pleistocene.

In July 2018, a team of Russian scientists in collaboration with Princeton University announced that they had brought two female nematodes frozen in permafrost, from around 42,000 years ago, back to life. The two nematodes, at the time, were the oldest confirmed living animals on the planet.^[30]^[31]

Pleistocene of Northern Spain, including woolly mammoth, cave lions eating a reindeer, horses, and woolly rhinoceros

Pleistocene of South America, including Megatherium and two Glyptodon

Humans

The evolution of anatomically modern humans took place during the Pleistocene.^[32]^[33] In the beginning of the Pleistocene Paranthropus species were still present, as well as early human ancestors, but during the lower Palaeolithic they disappeared, and the only hominin species found in fossilic records is Homo erectus for much of the Pleistocene. Acheulean lithics appear along with Homo erectus, some 1.8 million years ago, replacing the more primitive Oldowan industry used by A. garhi and by the earliest species of Homo. The Middle Paleolithic saw more varied speciation within Homo, including the appearance of Homo sapiens about 300,000 years ago.^[34]

According to mitochondrial timing techniques, modern humans migrated from Africa after the Riss glaciation in the Middle Palaeolithic during the Eemian Stage, spreading all over the ice-free world during the late Pleistocene.^[35]^[36]^[37] A 2005 study posits that humans in this migration interbred with archaic human forms already outside of Africa by the late Pleistocene, incorporating archaic human genetic material into the modern human gene pool.^[38]

**Hominin species during Pleistocene**

Explanatory notes

^ The Upper Pleistocene is a subseries/subepoch rather than a stage/age but, in 2009, the IUGS decided that it will be replaced with a stage/age (currently unofficially/informally named the Tarantian).^[15]

References

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^ Shatilovich, A. V.; Tchesunov, A. V.; Neretina, T. V.; Grabarnik, I. P.; Gubin, S. V.; Vishnivetskaya, T. A.; Onstott, T. C.; Rivkina, E. M. (16 July 2018). "Viable Nematodes from Late Pleistocene Permafrost of the Kolyma River Lowland". Doklady Biological Sciences. 480 (1): 100–102. doi:10.1134/S0012496618030079. PMID 30009350. S2CID 49743808.
^ Rogers, A.R.; Jorde, L.B. (1995). "Genetic evidence on modern human origins". Human Biology. 67 (1): 1–36. JSTOR 41465052. PMID 7721272.
^ Wall, J.D.; Przeworski, M. (2000). "When did the human population start increasing?". Genetics. 155 (4): 1865–1874. doi:10.1093/genetics/155.4.1865. PMC 1461207. PMID 10924481.
^ Hublin, Jean-Jacques; Ben-Ncer, Abdelouahed; Bailey, Shara E.; Freidline, Sarah E.; Neubauer, Simon; Skinner, Matthew M.; Bergmann, Inga; Le Cabec, Adeline; Benazzi, Stefano; Harvati, Katerina; Gunz, Philipp (8 June 2017). "New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens" (PDF). Nature. 546 (7657): 289–292. Bibcode:2017Natur.546..289H. doi:10.1038/nature22336. PMID 28593953.
^ Cann, R.L.; Stoneking, M.; Wilson, A.C. (1 January 1987). "Mitochondrial DNA and human evolution". Nature. 325 (6099): 31–36. Bibcode:1987Natur.325...31C. doi:10.1038/325031a0. PMID 3025745. S2CID 4285418.
^ Stringer, C.B. (1992) "Evolution of early modern humans" In: Jones, Steve; Martin, R. and Pilbeam, David R. (eds.) (1992) The Cambridge encyclopedia of human evolution Cambridge University Press, Cambridge, ISBN 0-521-32370-3, pp. 241–251.
^ Templeton, A. R. (7 March 2002). "Out of Africa again and again" (PDF). Nature. 416 (6876): 45–51. Bibcode:2002Natur.416...45T. doi:10.1038/416045a. PMID 11882887. S2CID 4397398. Archived from the original (PDF) on 12 April 2020. Retrieved 17 November 2017.
^ Eswarana, Vinayak; Harpendingb, Henry; Rogers, Alan R (July 2005). "Genomics refutes an exclusively African origin of humans". Journal of Human Evolution. 49 (1): 1–18. doi:10.1016/j.jhevol.2005.02.006. PMID 15878780.

Ogg, Jim (June 2004). "Overview of Global Boundary Stratotype Sections and Points (GSSP's)". International Commission on Stratigraphy. Retrieved 20 March 2019.

External links

[note_A-16] The Upper Pleistocene is a subseries/subepoch rather than a stage/age but, in 2009, the IUGS decided that it will be replaced with a stage/age (currently unofficially/informally named the Tarantian).^[15]

[ICC-1] Cohen, K. M.; Finney, S. C.; Gibbard, P. L.; Fan, J.-X. (January 2020). "International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy. Retrieved 23 February 2020.

[FRS-2] Mike Walker; et al. (December 2018). "Formal ratification of the subdivision of the Holocene Series/Epoch (Quaternary System/Period)" (PDF). Episodes. Subcommission on Quaternary Stratigraphy (SQS). 41 (4): 213–223. doi:10.18814/epiiugs/2018/018016. Retrieved 11 November 2019.

[3] Gibbard, Philip; Head, Martin (September 2010). "The newly-ratified definition of the Quaternary System/Period and redefinition of the Pleistocene Series/Epoch, and comparison of proposals advanced prior to formal ratification" (PDF). Episodes. 33 (3): 152–158. doi:10.18814/epiiugs/2010/v33i3/002. Retrieved 8 December 2020.

[4] Walker, Mike; Johnse, Sigfus; Rasmussen, Sune; Steffensen, Jørgen-Peder; Popp, Trevor; Gibbard, Phillip; et al. (June 2008). "The Global Stratotype Section and Point (GSSP) for the base of the Holocene Series/Epoch (Quaternary System/Period) in the NGRIP ice core". Episodes. 31 (2): 264–267. doi:10.18814/epiiugs/2008/v31i2/016.

[5] Jones, Daniel (2003) [1917], Peter Roach; James Hartman; Jane Setter (eds.), English Pronouncing Dictionary, Cambridge: Cambridge University Press, ISBN 3-12-539683-2

[6] Zalloua, Pierre A.; Matisoo-Smith, Elizabeth (6 January 2017). "Mapping Post-Glacial expansions: The Peopling of Southwest Asia". Scientific Reports. 7: 40338. Bibcode:2017NatSR...740338P. doi:10.1038/srep40338. ISSN 2045-2322. PMC 5216412. PMID 28059138.

[7] Lyell, Charles (1839). Nouveaux éléments de géologie (in French). Paris, France: Pitois-Levranet. p. 621. From p. 621: "Toutefois, en même temps … et de substituer à la dénomination de Nouveau Pliocène celle plus abrégée de Pleistocène, tirée du grec pleiston, plus, et kainos, récent." (However, at the same time that it became necessary to subdivide the two periods mentioned above, I found that the terms intended to designate these subdivisions were of an inconvenient length, and I have proposed to use in the future the word "Pliocene" for "old Pliocene", and to substitute for the name "new Pliocene" this shorter "Pleistocene", drawn from the Greek pleiston (most) and kainos (recent).)

[8] Wilmarth, Mary Grace (1925). Bulletin 769: The Geologic Time Classification of the United States Geological Survey Compared With Other Classifications, accompanied by the original definitions of era, period and epoch terms. Washington, D.C., U.S.A.: U.S. Government Printing Office. p. 47.

[OnlineEtDict-9] "Pleistocene". Online Etymology Dictionary.

[ICS_majordiv-10] Jump up to:^a ^b "Major Divisions". Subcommission on Quaternary Stratigraphy. International Commission on Stratigraphy. Retrieved 25 September 2019.

[11] For the top of the series, see: Lourens, L.; Hilgen, F.; Shackleton, N. J.; Laskar, J.; Wilson, D. (2004). "The Neogene Period". In Gradstein, F.; Ogg, J.; Smith, A. G. (eds.). A Geologic Time Scale 2004. Cambridge: Cambridge University Press. ISBN 0-521-78142-6.

[deBlij2012-12] Blij, Harm (2012). "Holocene Humanity". Why Geography Matters: More Than Ever (2nd ed.). Oxford: Oxford University Press. ISBN 978-0-19-991374-9.

[ICSChart1-13] "International Chronostratigraphic Chart v2017/02". International Commission on Stratigraphy. 2017. Retrieved 17 March 2018.

[Chiba1-14] "Japan-based name 'Chibanian' set to represent geologic age of last magnetic shift". The Japan Times. 14 November 2017. Retrieved 17 March 2018.

[ICSSQS-15] "Formal subdivision of the Pleistocene Series/Epoch". Subcommission on Quaternary Stratigraphy (International Commission on Stratigraphy). 4 January 2016. Retrieved 17 March 2018.

[17] Riccardi, Alberto C. (30 June 2009). "IUGS ratified ICS Recommendation on redefinition of Pleistocene and formal definition of base of Quaternary" International Union of Geological Sciences

[18] Svensson, A.; Nielsen, S. W.; Kipfstuhl, S.; Johnsen, S. J.; Steffensen, J. P.; Bigler, M.; Ruth, U.; Röthlisberger, R. (2005). "Visual stratigraphy of the North Greenland Ice Core Project (NorthGRIP) ice core during the last glacial period" (PDF). Journal of Geophysical Research. 110 (D2): D02108. Bibcode:2005JGRD..110.2108S. doi:10.1029/2004jd005134.

[Gradstein-28-19] Gradstein, Felix M.; Ogg, James G. and Smith, A. Gilbert (eds.) (2005) A Geologic Time Scale 2004 Cambridge University Press, Cambridge, UK, p. 28, ISBN 0-521-78142-6

[Rio-20] Rio, D.; Sprovieri, R.; Castradori, D.; Di Stefano, E. (1998). "The Gelasian Stage (Upper Pliocene): a new unit of the global standard chronostratigraphic scale" (PDF). Episodes. 21 (2): 82–87. doi:10.18814/epiiugs/1998/v21i2/002.

[21] Coles, J. M. (1975). The archaeology of early man. E. S. Higgs. Harmondsworth: Penguin. ISBN 0-14-055107-7. OCLC 2966860.

[22] National Geographic Channel, Six Degrees Could Change The World, Mark Lynas interview. Retrieved 14 February 2008.

[23] Yusuke Suganuma, Hideki Miura, Albert Zondervan, Jun'ichi Okuno (August 2014). "East Antarctic deglaciation and the link to global cooling during the Quaternary: evidence from glacial geomorphology and 10Be surface exposure dating of the Sør Rondane Mountains, Dronning Maud Land". Quaternary Science Reviews. 97: 102–120. Bibcode:2014QSRv...97..102S. doi:10.1016/j.quascirev.2014.05.007.

[24] "Quaternary Period". National Geographic. 6 January 2017.

[RichmondOther1-25] Jump up to:^a ^b Richmond, G.M.; Fullerton, D.S. (1986). "Summation of Quaternary glaciations in the United States of America". Quaternary Science Reviews. 5: 183–196. Bibcode:1986QSRv....5..183R. doi:10.1016/0277-3791(86)90184-8.

[RoyOther20041-26] Roy, M.; Clark, P.U.; Barendregt, R.W.; Glasmann; Enkin, R.J. (2004). "Glacial stratigraphy and paleomagnetism of late Cenozoic deposits of the north-central United States" (PDF). Geological Society of America Bulletin. 116 (1–2): 30–41. Bibcode:2004GSAB..116...30R. doi:10.1130/B25325.1. Archived from the original (PDF) on 28 September 2018. Retrieved 20 March 2010.

[27] Aber, J. S. (December 1991). "The Glaciation of Northeastern Kansas". Boreas. 20 (4): 297–314. doi:10.1111/j.1502-3885.1991.tb00282.x. (contains a summary of how and why the Nebraskan, Aftonian, Kansan, and Yarmouthian stages were abandoned by modern stratigraphers).

[Willeit-28] Willeit, M.; Ganopolski, A.; Calov, R.; Brovkin, V. (2019). "Mid-Pleistocene transition in glacial cycles explained by declining CO2 and regolith removal | Science Advances". Science Advances. 5 (4): eaav7337. doi:10.1126/sciadv.aav7337. PMC 6447376. PMID 30949580.

[29] Petra Bajo; et al. (2020). "Persistent influence of obliquity on ice age terminations since the Middle Pleistocene transition". Science. Vol. 367, no. 6483. pp. 1235–1239. doi:10.1126/science.aaw1114.

[30] Valdez, Raul. Mountain Sheep of North America.

[31] "Worms frozen in permafrost for up to 42,000 years come back to life". The Siberian Times. 26 July 2018. Retrieved 25 August 2021.

[32] Shatilovich, A. V.; Tchesunov, A. V.; Neretina, T. V.; Grabarnik, I. P.; Gubin, S. V.; Vishnivetskaya, T. A.; Onstott, T. C.; Rivkina, E. M. (16 July 2018). "Viable Nematodes from Late Pleistocene Permafrost of the Kolyma River Lowland". Doklady Biological Sciences. 480 (1): 100–102. doi:10.1134/S0012496618030079. PMID 30009350. S2CID 49743808.

[33] Rogers, A.R.; Jorde, L.B. (1995). "Genetic evidence on modern human origins". Human Biology. 67 (1): 1–36. JSTOR 41465052. PMID 7721272.

[34] Wall, J.D.; Przeworski, M. (2000). "When did the human population start increasing?". Genetics. 155 (4): 1865–1874. doi:10.1093/genetics/155.4.1865. PMC 1461207. PMID 10924481.

[35] Hublin, Jean-Jacques; Ben-Ncer, Abdelouahed; Bailey, Shara E.; Freidline, Sarah E.; Neubauer, Simon; Skinner, Matthew M.; Bergmann, Inga; Le Cabec, Adeline; Benazzi, Stefano; Harvati, Katerina; Gunz, Philipp (8 June 2017). "New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens" (PDF). Nature. 546 (7657): 289–292. Bibcode:2017Natur.546..289H. doi:10.1038/nature22336. PMID 28593953.

[36] Cann, R.L.; Stoneking, M.; Wilson, A.C. (1 January 1987). "Mitochondrial DNA and human evolution". Nature. 325 (6099): 31–36. Bibcode:1987Natur.325...31C. doi:10.1038/325031a0. PMID 3025745. S2CID 4285418.

[37] Stringer, C.B. (1992) "Evolution of early modern humans" In: Jones, Steve; Martin, R. and Pilbeam, David R. (eds.) (1992) The Cambridge encyclopedia of human evolution Cambridge University Press, Cambridge, ISBN 0-521-32370-3, pp. 241–251.

[38] Templeton, A. R. (7 March 2002). "Out of Africa again and again" (PDF). Nature. 416 (6876): 45–51. Bibcode:2002Natur.416...45T. doi:10.1038/416045a. PMID 11882887. S2CID 4397398. Archived from the original (PDF) on 12 April 2020. Retrieved 17 November 2017.

[39] Eswarana, Vinayak; Harpendingb, Henry; Rogers, Alan R (July 2005). "Genomics refutes an exclusively African origin of humans". Journal of Human Evolution. 49 (1): 1–18. doi:10.1016/j.jhevol.2005.02.006. PMID 15878780.

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